1GlaxoSmithKline Research and Development, King of Prussia, Pennsylvania; 2Channing Laboratory, Brigham and Women's Hospital and Harvard Medical School and 3Division of Pulmonary and Critical Care Medicine, Department of Medicine, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts; 4GlaxoSmithKline Research and Development, Research Triangle Park, North Carolina; 5Department of Radiology, University of British Columbia, Vancouver, British Columbia, Canada; 6University of Iowa, Iowa City, Iowa; 7Haukeland University Hospital and Institute of Medicine, University of Bergen, Bergen, Norway; and 8Cambridge Institute for Medical Research, University of Cambridge, Cambridge, United Kingdom
